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28. Hayashi, S., Nakajima, Y., Tanaka, Y., Breeden B.T. III, Kanazawa, Y., and Tsogtbaatar, C., in press. A hadrosauroid vertebra from the Upper Cretaceous Izumi Group, Kagawa Prefecture, Japan. Paleontological Research, in press.


27. Zakharov, Y.D., , Nakajima, Y.,  Arkhangelsky, M. S., Popov, A. M., Bondarenko, L. G., Pokrovsky, V. K., in press. New findings of Triassic marine vertebrates in Eastern Russia: ammonoid age control and evidence for ichthyopterygian affinities. Stratigraphy and Geological Correlation, in press.

26. 中島保寿,三上智之,泉 賢太郎,2024. パレオパラドキシア瑞浪釜戸標本に付随する骨片様化石についての予察的報告. 瑞浪市化石博物館研究報告 50(3), 51-57.

25. Li, Q., Liu, J., Klein, N., Nakajima, Y., Sander, P. M., 2023. Puberty in a Mesozoic reptile. Current Biology 33, 011-3016.e3.

24. Ishida, Y., Tagiri, M., Kato, T., Tsunoda, S., Nakajima, Y., Thuy, B., Numberger-Thuy L. D., and Fujita, T., 2023. The new brittle-star species Stegophiura takaisoensis (Echinodermata, Ophiuroidea) from the Pliocene of Ibaraki Prefecture, central Japan. Paleontological Research 28(1); 82-96

23. 加藤太一, 増川玄哉, 新山颯大, 中島保寿, 薗田哲平, 安藤寿男, 2022. 那珂湊層群から産出したスッポン類の肋板化石の再記載:北アメリカおよび中央アジアの大型スッポン類化石との比較. 茨城県自然博物館研究報告 (25):1-11

22. Nakajima, Y., Shigeta, Y., Houssaye, A., Zakharov, Y. D., Popov, A. M., Sander, P. M., 2022. Early Triassic ichthyopterygian fossils from the Russian Far East. Scientific Reports 12, 5546

21. Miyata, S., Yabumoto, Y., Nakajima, Y., Ito, Y. and Sasaki, T., 2022. A second specimen of the crossognathiform fish Apsopelix miyazakii from the Cretaceous Yezo Group of Mikasa area, central Hokkaido, Japan. Paleontological Research 26: 213-223.

20. Kanno, S., Tokumaru, S., Nakagaki, S., Nakajima, Y., Misaki, A., Hikida, Y., & Sato, T., 2022. Santonian-Campanian neoselachian faunas of the Upper Cretaceous Yezo Group in Nakagawa Town, Hokkaido, Japan. Cretaceous Research 105139.

19. Kato, T., Nakajima, Y.Shiseki, K. and Ando, H., 2021. Advanced mosasaurs from the Upper Cretaceous Nakaminato Group in Japan. Island Arc 30.1: e12431.

18. 中島保寿,三宅幸雄,田中猛,甲能直樹, 2021. 岐阜県高山市丹生川町旗鉾に分布する美濃帯ペルム系石灰岩から産出した骨化石.化石 109: 1-2

17. 加藤太一,薗田哲平,中島保寿,安藤寿男, 2020.上部白亜系那珂湊層群から産出した“翼竜類”(ヒタチナカリュウ) 標本の分類学的再検討.茨城県自然博物館研究報告 (23): 43-50

16. 宮田真也,平山 廉,中島保寿,前川 優,大倉正敏,佐々木猛智,2019. 岩手県久慈市の上部白亜系久慈層群玉川層より産出した板鰓類化石群の予察的検討.化石研究会会誌, vol.51 (2), pp.68-75.

15. Houssaye, A., Sander, P. M., Nakajima, Y., 2018. Structural, functional, and physiological signals in ichthyosaur vertebral centrum microanatomy and histology. Geodiversitas 42(2), 161-170.

14. Wintrich, T., Hayashi, S., Houssaye, A., Nakajima, Y., Sander, P. M., 2017. A Triassic plesiosaurian skeleton and bone histology inform on evolution of a unique body plan. Science Advances 3(12), e1701144

13. Nakajima, Y., Danilov, I. G., Hirayama, R., Sonoda, T. and Scheyer, T. M., 2017. Morphological and histological evidence for the oldest known softshell turtles from Japan. Journal of Vertebrate Paleontology, in press

12. Vitek, N., Danilov, I. G., Nakajima, Y. and Hirayama, R. 2017. CT-based redescription and improved phylogenetic taxon sampling of Cretaceous and Paleogene soft-shelled turtles (Trionychidae) of Asia, including the oldest crown trionychids. Journal of Systematic Paleontology 16(3), 199–211.

11. 中島保寿,2017.古生態復元に新たな視点をもたらすカメの骨組織研究.化石研究会会誌,50,p.22–27.

10. Kanno, S., Nakajima, Y., Hikida, Y. and Sato, T., 2017. Sphenodus (Chondrichthyes, Neoselachii) from the Upper Cretaceous in Nakagawa Town, Hokkaido, Japan. Paleontological Research 21(2), 122–130.

9. Shigeta, Y. and Nakajima, Y., 2017. Discovery of the early Spathian (late Olenekian, Early Triassic) ammonoid Tirolites in the Hiraiso Formation, South Kitakami, Northeast Japan. Paleontological Research 21, 37–43.

8. Nakajima, Y., Hirayama, R. and Endo, H. 2014. Turtle humeral microanatomy and its relationship to lifestyle. Biological Journal of Linnean Society, 113, p. 719–734.

7. Nakajima, Y., Houssaye, A. and Endo, H. 2014. Osteohistology of the ichthyopterygian reptile Utatsusaurus hataii (Reptilia: Ichthyopterygia): Implications for early ichthyosaur biology. Acta Paleontologica Polonica 59, 343–352.

6. Takahashi Y., Nakajima Y. and Sato, T. 2014. A new Early Triassic ichthyopterygian fossil from the Osawa Formation in Minamisanriku Town, Miyagi Prefecture, Japan. Paleontological Research 18, 258–262.

5. Nakajima, Y. and Izumi, K., 2014. Coprolites from the upper Osawa Formation (upper Spathian), northeastern Japan: Evidence for predation in a marine ecosystem 5 Myr after the end-Permian mass extinction. Palaeogeography, Palaeoclimatology, Palaeoecology 414, p. 225–232.

4. Hayashi, S., Houssaye, A., Nakajima, Y., Chiba, K., Ando, T., et al., 2013, Bone inner structure suggests increasing aquatic adaptations in Desmostylia (Mammalia, Afrotheria). PLoS ONE vol. 8: e59146. doi:10.1371/journal.pone.0059146

3. Nakajima, Y. and Endo, H., 2013. Comparative humeral microanatomy of terrestrial, semiaquatic, and aquatic carnivorans using micro–focus CT scan. Mammal Study 38, p. 1–8. (Mammal Study Award)

2. Nakajima, Y. and Schoch, R. R., 2011. The first temnospondyl amphibian from Japan. Journal of Vertebrate Paleontology 31, p. 1154–1157.

1. 中島保寿,櫻井和彦,平山 廉,2011.むかわ町立穂別博物館が所蔵するカメ化石.穂別博物館研究報告,26,p. 1–34.


2. 中島保寿,2016. ウミガメの捕食 ―海ののんびりハンターたち― p.100-103 in 国立科学博物館 特別展「海のハンター展」公式図録 pp.128

1. 中島保寿,2016. 糞化石からわかる大量絶滅後の世界 p. 40 in 名古屋市科学館 特別展「恐竜・化石研究所」公式ガイドブックpp. 80


6. 中島保寿, 2017. 世界最古のスッポンを福井県で発見 – バラバラの化石片から証拠を得るには. academist Journal,

5. 郡司芽久,栗原 望,川田伸一郎,中島保寿,和田直己,藤原慎一,2015.体骨格の適応進化を考える ―分類学・機能形態学・古生物学の観点から― 哺乳類科学,55 (1),p.108-109.

4. 中島保寿,泉 健太郎,2015. 糞化石から見えた大量絶滅直後の海の生態系.東京大学理学系研究科・理学部ニュース 46(5),p. 9 2015年1月.

3. 西澤真樹子(文・イラスト),藻谷亮介,中島保寿(監修), 2013.NPO法人大阪自然史センター編:南三陸化石ものしりブック②魚竜編—ギョリュウってどんな生きもの?— 南三陸ネイチャーセンター友の会.pp. 1-8. 2013年8 月.

2. 中島保寿, 2012.“The 1st International Symposium on Paleohistology”参加報告.化石vol. 91, p. 47-49.

1. 遠藤秀紀,中島保寿,林 昭次, 2010. かたちの学校14 骨内部構造からみる生態およびその進化.哺乳類科学 50, p.87-89.




16. Nakajima, Y.,, 2017. Reseaches in the Sanriku Fukkō National Park, Japan and our expectations for National Geological Park of Guanling. The International Symposium of Guanling Biota Protection and Featured Regional Touristic Development, Guanling, China (招待講演)

15. Nakajima, Y., Takahashi, S., Sasaki, O., Ehiro, M., Misaki, M. 2017. A preliminary report on the fossil fish assemblage from the Osawa Formation (Spathian, Lower Triassic) of Northeastern Japan. IGCP 630 Annual Meeting, 06. 14-16 (16) Sendai, Japan.


14. 中島保寿,2017. 中生代羊膜類の骨格に起きた革新的進化(Innovative evolutions in the skeleton of Mesozoic amniotes).JpGU-AGU Joint Meeting 2017. 千葉県幕張市,5/20~24


13. Sander, P. M., Hayashi, S., Houssaye, A., Nakajima, Y., Sato, T., and Wintrich, T., 2016. The independent evolution of endothermy in Plesiosauria: Evidence from their unique long bone histology. 76th Annual Meeting, Society of Vertebrate Paleontology, 29 of 26-29 October, Salt Lake City, Utah, USA

12. Sander, P. M., Wintrich, T., Hayashi, S., Houssaye, A., Nakajima, Y. and Sato, T., 2016. The evolution of a unique body plan: new evidence from a Triassic plesiosaur skeleton and long bone histology. XIV Annual Meeting of The European Association of Vertebrate Palaeontologists, 6-10 July, (p.171A in Programme and Abstract Book) Haarlem, Netherlands


11. Sander, P. M., Hayashi, S., Houssaye, A., Nakajima, Y., Sato, T. and Wintrich, T., 2015b. The evolution of plesiosaur bone histology: evidence from long bones and vertebrae. 75th Annual Meeting, Society of Vertebrate Paleontology, 14-17 October, Dallas, Texas, USA

10. Sander, P. M., Hayashi, S., Houssaye, A., Nakajima, Y., Sato, T. and Wintrich, T., 2015a. The evolution of plesiosaur bone histology: new evidence from new finds. 3rd International Symposium on Paleohistology, 2-5 July, Bonn, Germany2015

9. Takahashi, Y., Nakajima, Y. and Sato, T. 2014. An Early Triassic ichthyopterygian reptile from Miyagi, Japan. The 7th China- Korea-Japan Graduate Student Forum, China University of Geosciences, Beijing, China. September 26-29, 2014, 3rd Prize受賞

8. Nakajima, Y., Schoch, R. R., Klug, S., Takahashi, Y., Ogishima, M. and Sato, T. 2014. Life before dinosaurs in Japan: Recent findings of vertebrate fossils from the Triassic of South Kitakami Terrane. 1st International Symposium on Asian Dinosaurs. March 21-23, Fukui, Japan

7. Nakajima, Y., 2013. Estimation of the bone growth center using inner bone structural features and its application for paleohistology. 73rd Annual Meeting of the Society of Vertebrate Paleontology. , October 30-November 2, Los Angeles, California, USA

6. Nakajima, Y., Hirayama, R. and Endo, H. 2013. Bone microanatomy of turtle limb bones: Advances in paleobiology of testudinates. 10th International Congress of Vertebrate Morphology, Barcelona, Spain, 8-12 July(招待講演)

5. Hayashi, S. Nakajima, Y., Chiba, K., Sawamura, H. and Ando, T., 2011. Bone histology suggests increasing aquatic adaptations in Desmostylia. 71st Annual Meeting of the Society of Vertebrate Paleontology, November 2-5, Las Vegas, Nevada.

4. Nakajima, Y., 2011 The internal long bone microanatomy of terrestrial and aquatic turtles. First International Symposium on Paleohistology. July 18-20, Barcelona, Spain.

3. Hayashi, S., Nakajima, Y., Chiba, K., Inuzuka, N., Sawamura, H., Ando, T., Kaneko, N. and Osaki, T., 2010. Bone histological variations of desmostylians (Mammalia, Tethytheria). The Annual Symposium of Vertebrate Palaeontology and Comparative Anatomy 2010. 14-18 September, Cambridge, UK.

2. Nakaijma, Y., 2010. Evaluating the utility of limb bone internal structure as an indicator for aquatic adaptation of Testudines. The Annual Symposium of Vertebrate Palaeontology and Comparative Anatomy 2010. 14-18, September, Cambridge, UK

1. Nakajima, Y., Danilov, I. G. and Hirayama, R., 2009. Plywood-like structures in costal bones of mid- to late Cretaceous trionychids from Asia. Gaffney Turtle Symposium 09, Alberta, 16-19 September.









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